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The R groups that can be protonated/deprotonated include Lys, Arg, and His, which have a + 1 charge when protonated, and Glu and Asp (carboxylic acids), Tyr and Ser (alcohols), and Cys (thiol), which have zero charges when protonated

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[Phosphorylation and activation of glutathione peroxidase and… [PMID: 3698805] Laboratory literature summary: [Phosphorylation and activation of glutathione peroxidase and glutathione-S-transferase by cAMP-dependent protein kinase]